The Pomarine Skua Stercorarius pomarinus breeds in high-latitude arctic and subarctic areas of the USSR, passing through the North Sea and temperate North Atlantic en route to wintering areas in the Atlantic, generally north of the Equator (Wynne-Edwards in Bannerman 1963; Wallace & Bourne 1981; Cramp & Simmons 1983). Historically, this was the predominant species of late-au

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