Gulls Larus have a very diverse feeding ecology, but vertebrates form a significant part of the diet of some species (Tinbergen 1953). They are well known for their depredations on the eggs and young of colonial seabirds; they also attack and kill injured or sick birds as large as Curlew Numenius arquata (Tinbergen 1953) and American Wigeon Anas americana (Presnall 1968), and congeners such as Lit


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