There are many people who assisted me in the preparation of this paper. Firstly, for all their help with regards to specimens in their care I would like to thank Mark Adams, Joanne Cooper, Alison Harding, Catherine O’Carroll, Robert Prŷs-Jones and Hein van Grouw at the Natural History Museum, Tring; Jennifer Gallichan at the National Museum of Wales; Rachael Utting at t


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